Crosbycus dasycnemus composite
Crosbycus dasycnemus, composite from many sources







Crosbycus Roewer, 1914 is a monotypic genus of Taracidae[1].

Its type species Crosbycus dasycnemus (Crosby, 1911) occurs in China [2] and British Columbia (Canada), being one of the rarest intercontinental genera of harvestmen. All other Old World Crosbycus are juvenile Nemastomatidae[3][4]


Crosbycus in honor to arachnologist Cyrus Richard Crosby (1879–1937). Gender masculine.


Crosbycus originally in Nemastomatidae. When Kratochvíl divided this in subfamilies, he kept Crosbycus in Nemastomatinae [5]. Gruber excluded Crosbycus definitively from Nemastomatidae[6], without precise familial placement (“in Ischyropsalididae?”). Martens suggested, but did not formally describe a monotypic subfamily in Ischyropsalidoidea for Crosbycus[7][8]. Shear transferred Crosbycus to Ceratolasmatidae[9], placing it as sister to Hesperonemastoma. Giribet and collaborators included Hesperonemastoma in the Sabaconidae, but left Crosbycus unassigned to any family[10]. Schönhofer in his catalog [1] described Taracidae composed by Crosbycus, Hesperonemastoma and Taracus.


  1. 1.0 1.1 Schönhofer A.L. (2013) A taxonomic catalogue of the Dyspnoi Hansen and Sørensen, 1904 (Arachnida: Opiliones). Zootaxa, 3679(1), 1–68.
  2. Tsurusaki, N. & Song, Da-xiang (1993a) Occurrence of Crosbycus dasycnemus (Crosby) (Opiliones, Palpatores, Ceratolasmatidae) in China. Japanese Journal of Entomology, 61(1), 175?176.
  3. Šilhavý, V. (1956) Sekáči — Opilionidea. Fauna ČSR, No. 7. Nakladatelství Československé Akademie Věd, Praha, 272 pp., 10 plates (in Czech)
  4. Gruber, J. & Martens, J. (1968) Morphologie, Systematik und Ökologie der Gattung Nemastoma C.L. Koch (s.str.) (Opiliones, Nemastomatidae). Senckenbergiana biologica, 49(2), 137–172.
  5. Kratochvíl, J. (1958) Höhlenweberknechte Bulgariens (Palpatores — Nemastomatidae) [Jeskynní sekáči Bulharska (Palpatores — Nemastomatidae)]. Práce Brněnské základny Československé akademie věd (= Acta Academiae Scientiarum Čechoslovenicae Basis Brunensis), Brno, 30(12), 523–576, figs 1–47.
  6. Gruber, J. (1970) Die "Nemastoma"-Arten Nordamerikas (Ischyropsalididae, Opiliones, Arachnida). Annalen des Naturhistorischen Museums in Wien, 74, 129–144.
  7. Martens, J. (1976) Genitalmorphologie, System und Phylogenie der Weberknechte (Arachnida: Opiliones). Entomologica Germanica, Stuttgart, 3(1/2), 51–68.
  8. Martens, J. (1978) Spinnentiere, Arachnida: Weberknechte, Opiliones. Die Tierwelt Deutschlands. Vol. 64. G. Fischer Verlag, Jena. 464 pages.
  9. Shear, W.A. (1986) A cladistic analysis of the opilionid superfamily Ischyropsalidoidea, with description of the new family Ceratolasmatidae, the new genus Acuclavella and four new species. American Museum Novitates, New York City, 2844, 1–29.
  10. Giribet, G., Vogt, L., Pérez, A., Sharma, P. & Kury, A. (2010) A multilocus approach to harvestmen phylogeny with emphasis on biogeography and the phylogeny of Laniatores. Cladistics, 26(4), 408–437.

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