Nemastomoides elaveris Thevenin, 1901
Nemastomoides elaveris Thevenin, 1901. Reconstruction from Pocock (1911).







Nemastomoididae Petrunkevitch, 1955 is a family of Dyspnoi with only one fossil genus, † Nemastomoides Thevenin, 1901.

Placement and compositionEdit

Originally included in Troguloidea. Dunlop (2007: 255)[1] considered them to be stem Phalangida. The type species comes from Carboniferous of France. A second species from Mazon Creek, Illinois, originally in Protopilio, was added by Dunlop (2004)[2].


Nemastomoides Thevenin, 1901Edit

  • Nemastomoides elaveris Thevenin, 1901 -- Coal Measures of Commentry, northern France. Upper Carboniferous.
  • Nemastomoides longipes (Petrunkevitch, 1913) -- USA, Illinois, Mazon Creek. Upper Carboniferous, Pennsylvanic, Lower Allegheny.


  1. Dunlop, J.A. (2007) Paleontology. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (Eds.), Harvestmen: the biology of the Opiliones. x + 597 pages. Harvard University Press, Cambridge and London, pp 247–265.
  2. Dunlop, J. A. (2004a) The enigmatic fossil arachnid Kustarachne tenuipes Scudder, 1890 is a harvestman. In: Samu, F. & Szinetár, C. (Eds.), European Arachnology 2002. Plant Protection Institute & Berzsenyi College, Budapest, pp. 17–25.

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