Gjellerupia penis colorized

Zalmoxis sp. from Papua, distal part of penis. Rutrum highlighted in pink. Photo copyright A.B. Kury.

The rutrum is a feature present in the penis of Zalmoxidae and related families. It was first cursorily defined by Kury (2003: 103) as a "spade-like plate". A more extense definition with illustration may be found in Kury & Pérez (2007: 245).